his research has contirbuted greatly ------our understanding of this disease 填介词

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his research has contirbuted greatly ------our understanding of this disease 填介词
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his research has contirbuted greatly ------our understanding of this disease 填介词
his research has contirbuted greatly ------our understanding of this disease 填介词

his research has contirbuted greatly ------our understanding of this disease 填介词
his research has contirbuted greatly --to----our understanding of this disease
他的研究对于理解这种疾病起到了很大的促进作用.

答案是to

这里to是介词,引导做出贡献的领域