此句中,现在分词causing的施动者是谁,是主句主语studies,还是从句主语sex hormones.怎么找分词的施动者Studies of both animals and humans have shown that sex hormones somehow affect the stress response,causing females under

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此句中,现在分词causing的施动者是谁,是主句主语studies,还是从句主语sex hormones.怎么找分词的施动者Studies of both animals and humans have shown that sex hormones somehow affect the stress response,causing females under
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此句中,现在分词causing的施动者是谁,是主句主语studies,还是从句主语sex hormones.怎么找分词的施动者Studies of both animals and humans have shown that sex hormones somehow affect the stress response,causing females under
此句中,现在分词causing的施动者是谁,是主句主语studies,还是从句主语sex hormones.怎么找分词的施动者
Studies of both animals and humans have shown that sex hormones somehow affect the stress response,causing females under stress to produce more of the trigger chemicals than do males.

此句中,现在分词causing的施动者是谁,是主句主语studies,还是从句主语sex hormones.怎么找分词的施动者Studies of both animals and humans have shown that sex hormones somehow affect the stress response,causing females under
是sex hormones ,从句子翻译中也可以看出来
找分词的施动者 a.可以联系上下文意思 b.你所找的那个施动者跟分词的关系主动-ing,被动-ed

是sex hormones
找起来就是重新组合句子
sex hormones caused the females ...