英语翻译He first understood the development of different plant species from developments in bird species that he observed in the island.

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英语翻译He first understood the development of different plant species from developments in bird species that he observed in the island.
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英语翻译He first understood the development of different plant species from developments in bird species that he observed in the island.
英语翻译
He first understood the development of different plant species from developments in bird species that he observed in the island.

英语翻译He first understood the development of different plant species from developments in bird species that he observed in the island.
He first understood the development of different plant species from developments in bird species that he observed in the island.
主语He,谓语understood,宾语the development,后面的(of different plant species)( from developments in bird species)( that he observed in the island)都是定语,其中第三个定语that he observed in the island修饰的是species

He(主语) first(定语) understood(谓语) the development(宾语) of different plant species(宾语) from developments in bird species(宾语补足语) that he observed in the island(定语从句修饰bird species).